香坂真一郎（长谷川博己饰）是搜查一课的刑警，能力相当出众，是公认的“未来的搜查一科科长”。香坂的父亲淳史一直想成为搜查一科科长却仍然停留在分局，香坂为了父亲、为了妻子美沙以及为了自己，向着搜查一科科长的目标迈进。同样想成为搜查一课课长的刑警山田春彦（冈田将生饰）从东京大学法学部毕业后便进入警视厅工作。他的父亲则是从警察厅次官晋升至内阁官房副长官的高层大人物。某日，香坂因犯了错误被调离总部，成为分局芝署的代理科长。他忍受着同事们的讥讽，努力想回到总部
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